Tired of dodging fishwives' brooms and carriage wheels, alley cat Skilley yearns for the comfort of Ye olde Cheshire Cheese, a popular London inn. When he hears that the innkeeper is in need of a mouser, he hatches a scheme so audacious it will surely set him up for the rest of his nine lives. Once installed at the Cheese, Skilley looks forward to a life of ease. But a resident mouse named Pip uncovers Skilley's scandalous secret, and the desperate cat is forced to make a pact with him. The two become allies, and harmony reigns until they are drawn into an intrigue involving a tyrannical cook, a strange visitor hidden in the attic, and an evil tomcat called Pinch. The ensuing mayhem threatens the peace of Ye olde Cheshire Cheese—and the entire British empire!